What is the meaning of the howling during the night?

From the text:

And when, on the still cold nights, he pointed his nose at a star and howled long and wolflike, it was his ancestors, dead and dust, pointing nose at star and howling down through the centuries and through him. And his cadences were their cadences, the cadences which voiced their woe and what to them was the meaning of the stiffness, and the cold, and dark.